Defender Tomkins, 33, joined the Eagles from West Ham in 2016 for £10m.

Ghana forward Ayew, 30, joined from Swansea in 2018, initially on loan, while youth product Clyne, 31, returned to the club in 2020.

The quartet have made a combined 677 appearances for Palace, with Tomkins contributing 124, Ayew 133 and right-back Clyne 171.

"It remains of pivotal importance to get the right balance of youth and experience in any Premier League squad," said Palace chairman Steve Parish.

"I'm delighted that we've been able to ensure that these long-standing and experienced players in Jordan, Nathaniel, James and James remain with us for the upcoming campaign.

"Each of them played an enormous role in the progress we've all enjoyed in the last year, and we very much expect similar contributions for the season ahead."

The club have not yet released their retained list but remain in discussions with Cheikhou Kouyate, with the midfielder's contract due to expire at the end of this month.